Howlongatimeitwouldrequirebeforesuchacrossedbodyofanimalswouldassumeauniformcharacterwithinalimitedarea,noonecansay;thattheywouldultimatelybecomeuniformfromfreeintercrossing,andfromthesurvivalofthefittest,wemayfeelassured;butthecharactersthusacquiredwouldrarelyornever,asmaybeinferredfromthepreviousconsiderations,beexactlyintermediatebetweenthoseofthetwoparent—breeds。Withrespecttotheveryslightdifferencesbywhichtheindividualsofthesamesub—variety,orevenofalliedvarieties,arecharacterised,itisobviousthatfreecrossingwouldsoonobliteratesuchsmalldistinctions。Theformationofnewvarieties,independentlyofselection,wouldalsothusbeprevented;exceptwhenthesamevariationcontinuallyrecurredfromtheactionofsomestronglypredisposingcause。Wemaythereforeconcludethatfreecrossinghasinallcasesplayedanimportantpartingivinguniformityofcharactertoallthemembersofthesamedomesticraceandofthesamenaturalspecies,thoughlargelygovernedbynaturalselectionandbythedirectactionofthesurroundingconditions。
ONTHEPOSSIBILITYOFALLORGANICBEINGSOCCASIONALLYINTERCROSSING。
Butitmaybeasked,canfreecrossingoccurwithhermaphroditeanimalsandplants?Allthehigheranimals,andthefewinsectswhichhavebeendomesticated,haveseparatesexes,andmustinevitablyuniteforeachbirth。
Withrespecttothecrossingofhermaphrodites,thesubjectistoolargeforthepresentvolume,butinthe’OriginofSpecies’Ihavegivenashortabstractofthereasonswhichinducemetobelievethatallorganicbeingsoccasionallycross,thoughperhapsinsomecasesonlyatlongintervalsoftime。(15/14。Withrespecttoplants,anadmirableessayonthissubject(DieGeschlechter—VertheilungbeidenPflanzen:1867)hasbeenpublishedbyDr。
HildebrandwhoarrivesatthesamegeneralconclusionsasIhavedone。Variousothertreatiseshavesinceappearedonthesamesubject,moreespeciallybyHermannMullerandDelpino。)Iwillmerelyrecallthefactthatmanyplants,thoughhermaphroditeinstructure,areunisexualinfunction;——suchasthosecalledbyC。K。SprengelDICHOGAMOUS,inwhichthepollenandstigmaofthesameflowerarematuredatdifferentperiods;orthosecalledbymeRECIPROCALLYDIMORPHIC,inwhichtheflower’sownpollenisnotfittedtofertiliseitsownstigma;oragain,themanykindsinwhichcuriousmechanicalcontrivancesexist,effectuallypreventingself—fertilisation。Thereare,however,manyhermaphroditeplantswhicharenotinanywayspeciallyconstructedtofavourintercrossing,butwhichneverthelesscomminglealmostasfreelyasanimalswithseparatedsexes。Thisisthecasewithcabbages,radishes,andonions,asIknowfromhavingexperimentedonthem:eventhepeasantsofLiguriasaythatcabbagesmustbeprevented\"fromfallinginlove\"