IcarefullyexaminedtheflowersofninetyplantsofthecrossedAntirrhinuminthetwobeds,andtheirstructurehadnotbeenintheleastaffectedbythecross,exceptthatinafewinstancestheminuterudimentofthefifthstamen,whichisalwayspresent,wasmorefullyorevencompletelydeveloped。Itmustnotbesupposedthatthisentireobliterationofthepeloricstructureinthecrossedplantscanbeaccountedforbyanyincapacityoftransmission;forI
raisedalargebedofplantsfromthepeloricAntirrhinum,artificiallyfertilisedbyitsownpollen,andsixteenplants,whichalonesurvivedthewinter,wereallasperfectlypeloricastheparent—plant。Herewehaveagoodinstanceofthewidedifferencebetweentheinheritanceofacharacterandthepoweroftransmittingittocrossedoffspring。Thecrossedplants,whichperfectlyresembledthecommonsnapdragon,wereallowedtosowthemselves,andoutofahundredandtwenty—sevenseedlings,eighty—eightprovedtobecommonsnapdragons,twowereinanintermediateconditionbetweenthepeloricandnormalstate,andthirty—sevenwereperfectlypeloric,havingrevertedtothestructureoftheironegrand—parent。Thiscaseseemsatfirstsighttoofferanexceptiontotherulejustgiven,namely,thatacharacterwhichispresentinoneformandlatentintheotherisgenerallytransmittedwithprepotentforcewhenthetwoformsarecrossed。ForinalltheScrophulariaceae,andespeciallyinthegeneraAntirrhinumandLinaria,thereis,aswasshowninthelastchapter,astronglatenttendencytobecomepeloric;butthereisalso,aswehaveseen,astillstrongertendencyinallpeloricplantstoreacquiretheirnormalirregularstructure。Sothatwehavetwoopposedlatenttendenciesinthesameplants。Now,withthecrossedAntirrhinumsthetendencytoproducenormalorirregularflowers,likethoseofthecommonSnapdragon,prevailedinthefirstgeneration;whilstthetendencytopelorism,appearingtogainstrengthbytheintermissionofageneration,prevailedtoalargeextentinthesecondsetofseedlings。Howitispossibleforacharactertogainstrengthbytheintermissionofageneration,willbeconsideredinthechapteronpangenesis。
Onthewhole,thesubjectofprepotencyisextremelyintricate,——fromitsvaryingsomuchinstrength,eveninregardtothesamecharacter,indifferentanimals,——fromitsrunningeitherequallyinbothsexes,or,asfrequentlyisthecasewithanimals,butnotwithplants,muchstrongerinonesexthantheother,——fromtheexistenceofsecondarysexualcharacters,——fromthetransmissionofcertaincharactersbeinglimited,asweshallimmediatelysee,bysex,——fromcertaincharactersnotblendingtogether,——and,perhaps,occasionallyfromtheeffectsofapreviousfertilisationonthemother。Itisthereforenotsurprisingthatnoonehashithertosucceededindrawingupgeneralrulesonthesubjectofprepotency。
INHERITANCEASLIMITEDBYSEX。
Newcharactersoftenappearinonesex,andareafterwardstransmittedtothesamesex,eitherexclusivelyorinamuchgreaterdegreethantotheother。