Anothercase，thoughsomewhatdifferent，maybeheregiven，asitishighlyremarkable，andisestablishedonexcellentevidence。Kolreuterminutelydescribesfivevarietiesofthecommontobacco（16/25。’ZweiteForts。’s。53
namelyNicotianamajorvulgaris;（2）perennis;（3）transylvanica;（4）asub—
var。ofthelast;（5）majorlatifol。fl。alb。）whichwerereciprocallycrossed，andtheoffspringwereintermediateincharacterandasfertileastheirparents:fromthisfactKolreuterinferredthattheyarereallyvarieties;andnoone，asfarasIcandiscover，seemstohavedoubtedthatsuchisthecase。HealsocrossedreciprocallythesefivevarietieswithN。
glutinosa，andtheyyieldedverysterilehybrids;butthoseraisedfromthevar。perennis，whetherusedasthefatherormotherplant，werenotsosterileasthehybridsfromthefourothervarieties。（16/26。KolreuterwassomuchstruckwiththisfactthathesuspectedthatalittlepollenofN。glutinosainoneofhisexperimentsmighthaveaccidentallygotmingledwiththatofvar。perennis，andthusaideditsfertilisingpower。ButwenowknowconclusivelyfromGartner（’Bastarderz。’s。34，43）thatthepollenoftwospeciesneveractsCONJOINTLYonathirdspecies;stilllesswillthepollenofadistinctspecies，mingledwithaplant’sownpollen，ifthelatterbepresentinsufficientquantity，haveanyeffect。Thesoleeffectofminglingtwokindsofpollenistoproduceinthesamecapsuleseedswhichyieldplants，sometakingaftertheoneandsomeaftertheotherparent。）Sothatthesexualcapacityofthisonevarietyhascertainlybeeninsomedegreemodified，soastoapproachinnaturethatofN。glutinosa。（16/27。Mr。Scotthasmadesomeobservationsontheabsolutesterilityofapurpleandwhiteprimrose（Primulavulgaris）whenfertilisedbypollenfromthecommonprimrose（’JournalofProc。ofLinn。Soc。’volume81864page98）;buttheseobservationsrequireconfirmation。Iraisedanumberofpurple—floweredlong—
styledseedlingsfromseedkindlysentmebyMr。Scott，and，thoughtheywereallinsomedegreesterile，theyweremuchmorefertilewithpollentakenfromthecommonprimrosethanwiththeirownpollen。Mr。Scotthaslikewisedescribedaredequal—styledcowslip（P。verisibidpage106），whichwasfoundbyhimtobehighlysterilewhencrossedwiththecommoncowslip;butthiswasnotthecasewithseveralequal—styledredseedlingsraisedbymefromhisplant。Thisvarietyofthecowslippresentstheremarkablepeculiarityofcombiningmaleorgansineveryrespectlikethoseoftheshort—styledform，withfemaleorgansresemblinginfunctionandpartlyinstructurethoseofthelong—styledform;sothatwehavethesingularanomalyofthetwoformscombinedinthesameflower。Henceitisnotsurprisingthattheseflowersshouldbespontaneouslyself—fertileinahighdegree。）