Takingallthesefactstogether,itisobviousthattheview,whichwasentertainedbyMantellandtheprobabilityofwhichwasdemonstratedbyyourowndistinguishedanatomist,Leidy,whilemuchadditionalevidenceinthesamedirectionhasbeenfurnishedbyProfessorCope,thatsomeoftheseanimalsmayhavewalkedupontheirhindlegsasbirdsdo,acquiresgreatweight。
Infact,therecanbenoreasonabledoubtthatoneofthesmallerformsoftheOrnithoscelida,Compsognathus,thealmostentireskeletonofwhichhasbeendiscoveredintheSolenhofenslates,wasabipedalanimal。Thepartsofthisskeletonaresomewhattwistedoutoftheirnaturalrelations,buttheaccompanyingfiguregivesajustviewofthegeneralformofCompsognathusandoftheproportionsofitslimbs;which,insomerespects,aremorecompletelybird-likethanthoseofotherOrnithoscelida。
Fig。7。——RestorationofCompsognathusLongipesWehavehadtostretchthedefinitionoftheclassofbirdssoastoincludebirdswithteethandbirdswithpaw-likeforelimbsandlongtails。ThereisnoevidencethatCompsognathuspossessedfeathers;but,ifitdid,itwouldbehardindeedtosaywhetheritshouldbecalledareptilianbirdoranavianreptile。
AsCompsognathuswalkeduponitshindlegs,itmusthavemadetrackslikethoseofbirds。AndasthestructureofthelimbsofseveralofthegiganticOrnithoscelida,suchasIguanodon,leadstotheconclusionthattheyalsomayhaveconstantly,oroccasionally,assumedthesameattitude,apeculiarinterestattachestothefactthat,intheWealdenstrataofEngland,therearetobefoundgiganticfootsteps,arrangedinorderlikethoseoftheBrontozoum,andwhichtherecanbenoreasonabledoubtweremadebysomeoftheOrnithoscelida,theremainsofwhicharefoundinthesamerocks。And,knowingthatreptilesthatwalkedupontheirhindlegsandsharedmanyoftheanatomicalcharactersofbirdsdidonceexist,itbecomesaveryimportantquestionwhetherthetracksintheTriasofMassachusetts,towhichIreferredsometimeago,andwhichformerlyusedtobeunhesitatinglyascribedtobirds,maynotallhavebeenmadebyornithoscelidanreptiles;andwhether,ifwecouldobtaintheskeletonsoftheanimalswhichmadethesetracks,weshouldnotfindinthemtheactualstepsoftheevolutionalprocessbywhichreptilesgaverisetobirds。
TheevidentialvalueofthefactsIhavebroughtforwardinthisLecturemustbeneitherovernorunderestimated。Itisnothistoricalproofoftheoccurrenceoftheevolutionofbirdsfromreptiles,forwehavenosafegroundforassumingthattruebirdshadnotmadetheirappearanceatthecommencementoftheMesozoicepoch。Itis,infact,quitepossiblethatallthesemoreorlessavi-formreptilesoftheMesozoicepochsarenottermsintheseriesofprogressionfrombirdstoreptilesatall,butsimplythemoreorlessmodifieddescendantsofPalaeozoicformsthroughwhichthattransitionwasactuallyeffected。