Havingnowendeavouredtoshowthattheseveralforegoingassumptionsaretoacertainextentsupportedbyanalogousfacts,andhavingalludedtosomeofthemostdoubtfulpoints,wewillconsiderhowfarthehypothesisbringsunderasinglepointofviewthevariouscasesenumeratedintheFirstPart。Alltheformsofreproductiongraduateintooneanotherandagreeintheirproduct;
foritisimpossibletodistinguishbetweenorganismsproducedfrombuds,fromself—division,orfromfertilisedgerms;suchorganismsareliabletovariationsofthesamenatureandtoreversionsofthesamekind;andas,accordingtoourhypothesis,alltheformsofreproductiondependontheaggregationofgemmulesderivedfromthewholebody,wecanunderstandthisremarkableagreement。Parthenogenesisisnolongerwonderful,andifwedidnotknowthatgreatgoodfollowedfromtheunionofthesexualelementsderivedfromtwodistinctindividuals,thewonderwouldbethatparthenogenesisdidnotoccurmuchoftenerthanitdoes。Onanyordinarytheoryofreproductiontheformationofgraft—hybrids,andtheactionofthemaleelementonthetissuesofthemother—plant,aswellasonthefutureprogenyoffemaleanimals,aregreatanomalies;buttheyareintelligibleonourhypothesis。Thereproductiveorgansdonotactuallycreatethesexualelements;theymerelydeterminetheaggregationandperhapsthemultiplicationofthegemmulesinaspecialmanner。Theseorgans,however,togetherwiththeiraccessoryparts,havehighfunctionstoperform。Theyadaptoneorbothelementsforindependenttemporaryexistence,andformutualunion。Thestigmaticsecretionactsonthepollenofaplantofthesamespeciesinawhollydifferentmannertowhatitdoesonthepollenofonebelongingtoadistinctgenusorfamily。ThespermatophoresoftheCephalopodaarewonderfullycomplexstructures,whichwereformerlymistakenforparasiticworms;andthespermatozoaofsomeanimalspossessattributeswhich,ifobservedinanindependentanimal,wouldbeputdowntoinstinctguidedbysense—organs,——aswhenthespermatozoaofaninsectfindtheirwayintotheminutemicropyleoftheegg。
Theantagonismwhichhaslongbeenobserved(27/55。Mr。HerbertSpencer(’PrinciplesofBiology’volume2page430)hasfullydiscussedthisantagonism。),withcertainexceptions,betweengrowthandthepowerofsexualreproduction(27/56。Themalesalmonisknowntobreedataveryearlyage。
TheTritonandSiredon,whilstretainingtheirlarvalbranchiae,accordingtoFilippiandDumeril(’AnnalsandMag。ofNat。Hist。’3rdseries1866page157)
arecapableofreproduction。ErnstHaeckelhasrecently(’MonatsberichtAkad。