The Variation of Animals and Plants under Domestic

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第35章

The Variation of Animals and Plants under Domestic Charles Darwin

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The Variation of Animals and Plants under Domestic
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  forinstance,somekindsofmossesandlichenshaveneverbeenseentofructifyinFrance。

  Someoftheseendemicandnaturalisedplantsareprobablyrenderedsterilefromexcessivemultiplicationbybuds,andtheirconsequentincapacitytoproduceandnourishseed。Butthesterilityofothersmoreprobablydependsonthepeculiarconditionsunderwhichtheylive,asinthecaseoftheivyinthenorthernpartofEurope,andofthetreesintheswampsoftheUnitedStates;yettheseplantsmustbeinsomerespectseminentlywelladaptedforthestationswhichtheyoccupy,fortheyholdtheirplacesagainstahostofcompetitors。]

  Finally,thehighdegreeofsterilitywhichoftenaccompaniesthedoublingofflowers,oranexcessivedevelopmentoffruit,seldomsupervenesatonce。Anincipienttendencyisobserved,andcontinuedselectioncompletestheresult。

  Theviewwhichseemsthemostprobable,andwhichconnectstogetheralltheforegoingfactsandbringsthemwithinourpresentsubject,is,thatchangedandunnaturalconditionsoflifefirstgiveatendencytosterility;andinconsequenceofthis,theorgansofreproductionbeingnolongerablefullytoperformtheirproperfunctions,asupplyoforganisedmatter,notrequiredforthedevelopmentoftheseed,flowseitherintotheseorgansandrendersthemfoliaceous,orintothefruit,stems,tubers,etc。,increasingtheirsizeandsucculency。Butitisprobablethatthereexists,independentlyofanyincipientsterility,anantagonismbetweenthetwoformsofreproduction,namely,byseedandbuds,wheneitheriscarriedtoanextremedegree。Thatincipientsterilityplaysanimportantpartinthedoublingofflowers,andintheothercasesjustspecified,Iinferchieflyfromthefollowingfacts。Whenfertilityislostfromawhollydifferentcause,namely,fromhybridism,thereisastrongtendency,asGartner(18/118。’Bastarderzeugung’s。565。Kolreuter’DritteFortsetzung’s。73,87,119)alsoshowsthatwhentwospecies,onesingleandtheotherdouble,arecrossed,thehybridsareapttobeextremelydouble。)affirms,forflowerstobecomedouble,andthistendencyisinherited。Moreover,itisnotoriousthatwithhybridsthemaleorgansbecomesterilebeforethefemaleorgans,andwithdoubleflowersthestamensfirstbecomefoliaceous。Thislatterfactiswellshownbythemaleflowersofdioeciousplants,which,accordingtoGallesio(18/119。’TeoriadellaRiproduzioneVeg。’1816page73。)firstbecomedouble。Again,Gartner(18/120。

  ’Bastarderzeugung’s。573。)ofteninsiststhattheflowersofevenutterlysterilehybrids,whichdonotproduceanyseed,generallyyieldperfectcapsulesorfruit,——afactwhichhaslikewisebeenrepeatedlyobservedbyNaudinwiththeCucurbitaceae;sothattheproductionoffruitbyplantsrenderedsterilethroughanycauseisintelligible。Kolreuterhasalsoexpressedhisunboundedastonishmentatthesizeanddevelopmentofthetubersincertainhybrids;andallexperimentalists(18/121。Ibids。527。)haveremarkedonthestrongtendencyinhybridstoincreasebyroots,runners,andsuckers。Seeingthathybridplants,whichfromtheirnaturearemoreorlesssterile,thustendtoproducedoubleflowers;thattheyhavethepartsincludingtheseed,thatisthefruit,perfectlydeveloped,evenwhencontainingnoseed;thattheysometimesyieldgiganticroots;thattheyalmostinvariablytendtoincreaselargelybysuckersandothersuchmeans;——seeingthis,andknowing,fromthemanyfactsgivenintheearlierpartsofthischapter,thatalmostallorganicbeingswhenexposedtounnaturalconditionstendtobecomemoreorlesssterile,itseemsmuchthemostprobableviewthatwithcultivatedplantssterilityistheexcitingcause,anddoubleflowers,richseedlessfruit,andinsomecaseslargely—developedorgansofvegetation,etc。,aretheindirectresults——theseresultshavingbeeninmostcaseslargelyincreasedthroughcontinuedselectionbyman。

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