MyworkontheCirripediapossesses,Ithink,considerablevalue,asbesidesdescribingseveralnewandremarkableforms,Imadeoutthehomologiesofthevariousparts——Idiscoveredthecementingapparatus,thoughIblundereddreadfullyaboutthecementglands——andlastlyIprovedtheexistenceincertaingeneraofminutemalescomplementaltoandparasiticonthehermaphrodites。Thislatterdiscoveryhasatlastbeenfullyconfirmed;thoughatonetimeaGermanwriterwaspleasedtoattributethewholeaccounttomyfertileimagination。TheCirripedesformahighlyvaryinganddifficultgroupofspeciestoclass;andmyworkwasofconsiderableusetome,whenIhadtodiscussinthe’OriginofSpecies’
theprinciplesofanaturalclassification。Nevertheless,Idoubtwhethertheworkwasworththeconsumptionofsomuchtime。
>FromSeptember1854Idevotedmywholetimetoarrangingmyhugepileofnotes,toobserving,andtoexperimentinginrelationtothetransmutationofspecies。Duringthevoyageofthe\"Beagle\"IhadbeendeeplyimpressedbydiscoveringinthePampeanformationgreatfossilanimalscoveredwitharmourlikethatontheexistingarmadillos;secondly,bythemannerinwhichcloselyalliedanimalsreplaceoneanotherinproceedingsouthwardsovertheContinent;andthirdly,bytheSouthAmericancharacterofmostoftheproductionsoftheGalapagosarchipelago,andmoreespeciallybythemannerinwhichtheydifferslightlyoneachislandofthegroup;noneoftheislandsappearingtobeveryancientinageologicalsense。
Itwasevidentthatsuchfactsasthese,aswellasmanyothers,couldonlybeexplainedonthesuppositionthatspeciesgraduallybecomemodified;andthesubjecthauntedme。Butitwasequallyevidentthatneithertheactionofthesurroundingconditions,northewilloftheorganisms(especiallyinthecaseofplants)couldaccountfortheinnumerablecasesinwhichorganismsofeverykindarebeautifullyadaptedtotheirhabitsoflife——
forinstance,awoodpeckeroratree—frogtoclimbtrees,oraseedfordispersalbyhooksorplumes。Ihadalwaysbeenmuchstruckbysuchadaptations,anduntilthesecouldbeexplaineditseemedtomealmostuselesstoendeavourtoprovebyindirectevidencethatspecieshavebeenmodified。